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The circRNAs were involved in signaling pathways that regulate stem cell pluripotency (e.g., AKT3, AVCVR2A, FGFR1, ACVR1, FZD3, SMAD4 ), tight junction ( MYH15, PRKCA, AMOTL1, PPP2CB, CDC42, PTEN ), and adhesion connections (e.g., LEF1, NECTIN3, SMAD2, AFDN ), hedgehog signaling pathways (e.g., HHIP, GSK3B, PTCH1, SMO, RAB23 ), cAMP signaling pathways (e.g., PLD1, TIAM1, GNAI1, ADCY1 ), mTOR signaling pathways (e.g., BRAF, RICTOR, HIF1A ), and phosphatidylinositol signaling systems (e.g., CDS1, DGKH, PLCB1, DGKK )