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Xu's study [60] found that Pio, a peroxisome proliferatoractivated receptor agonist, could induce apoptosis and compensate for this effect by activating autophagy in TSPCs, thereby improving TSPC dysfunction

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In addition, LKB1 inactivation, in conjunction with KRAS G12D mutation, promotes serine biosynthesis, contributing to DNA methylation and the enrichment at retrotransposon elements associated with their transcriptional silencing [102]