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For this reason, the homeostasis of extracellular glutamate is tightly controlled by the combined activity of neuronal and glial membrane transporters, such as glutamate transporter-1 (GLT1), glutamate aspartate transporter (GLAST), and the cysteine/glutamate antiporter, or system xc (Sxc ), present in astrocytes, and EAAT3 present in neurons

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It activates calcium-calmodulin mediated protein kinase beta (CaMKK), liver kinase B1 (LKB1), and upstream kinases (transforming growth factor--activated kinase 1 (TAK1)), that phosphorylates and activate AMPK